Bacopa

Scientific name

Bacopa Aubl.

Common names

water hyssop, herb of grace, brahmi

Family

Scrophulariaceae

Similar genera

Diodia, Lindernia, Lysimachia, Macbridea, Micranthemum, Myriophyllum (rarely confused), Oldenlandia, Rotala, Veronica

Native distribution

tropical and subtropical regions of world, particularly America

Species cultivated

Bacopa australis D.C. Souza

B. caroliniana (Walter) B.L. Rob. (B. amplexicaulis)

B. innominata (M.Gómez) Alain

B. lanigera (Cham. & Schldl.) Wettst.

B. monnieri (L.) Wettst.

B. myriophylloides (Benth.) Wettst.

B. salzmannii (Benth.) Edwall

B. serpyllifolia (Benth.) Pennell

Adventive distribution

Bacopa caroliniana is introduced into Australia.

B. egensis (Poepp.) Pennell is introduced into the United States.

Weed status

Native species of Bacopa (B. rotundifolia (Michx.) Wettst., B. monnieri, and B. eisenii (Kell.) Pennell) are known to be weeds in rice crops.

Habit

terrestrial, amphibiousamphibious:
(adj) of a plant able to live on land or in water
or obligately aquatic stem plant

Brief description

Annual or perennialperennial:
(adj) (of a plant) having a life cycle of more than two years
, decumbentdecumbent:
(adj) (of stems) having a portion lying along the ground, with upper parts erect or ascending
or erect. Stem hairy or smooth. Leaves oppositeopposite:
(adj) (of leaves) two leaves per node; in pairs on opposite sides of an axis
or whorledwhorled:
(n) bearing whorls; a type of leaf arrangement (phyllotaxis) in which leaves are in whorls
, sessilesessile:
(adj) attached directly, without a stalk
; leaf bladeblade:
(n) (syn. lamina) the flat, expanded part of a leaf, frond, or petal (excluding, e.g., the petiole)
round to linear, venationvenation:
(n) the arrangement of veins in a leaf
palmatepalmate:
(adj) (of leaves or venation) with lobes, leaflets, divisions or veins originating from the same point
or pinnatepinnate:
(adj) in the form of a feather; of, e.g., leaflets, lobes, or veins: arranged in two rows along an axis
. Flowers solitary or in pairs in leaf axils, usually actinomorphicactinomorphic:
(adj) of flowers, having radial symmetry; capable of being bisected into identifical halves along more than one axis
, with 5 sepals and 5 petals, usually white, blue or purple. Dispersal and propagation by seeds and stem fragments.

Natural habitat

often found as a littorallittoral:
(adj) of or along the shore
inhabitant of streams, lakes, and wetlands

Additional comments

This genus contains 70-100 species, most of which are terrestrialterrestrial:
(adj) growing on land as opposed to living in water
or amphibiousamphibious:
(adj) of a plant able to live on land or in water
; several species (e.g., B. myriophylloides) are obligate aquatic. Only four species are commonly cultivated. A terrestrialterrestrial:
(adj) growing on land as opposed to living in water
plant with coarsely toothed leaves and white flowers sold as Bacopa is actually Sutera diffusus. Species of Bacopa are usually aromatic, with a lemon scent when damaged, and plant extracts are used as herbal remedies.

Hydrotrida caroliniana (Walter) Small is a synonym of Bacopa caroliniana (Walter) B.L.Rob. and only distinguished by the presence of a ring of bristles around the base of the ovaryovary:
(n) a hollow organ at the base of the carpel of a flower in which ovules are produced
. One character is not sufficient enough to form a separate genus, especially when several other features are shared (Pennell, 1946). As in most literature and flora, we treat Bacopa caroliniana (Walter) B.L.Rob. as the accepted name.

  Bacopa australis , emersed; photo: S.L. Winterton

Bacopa australis, emersed; photo: S.L. Winterton

  Bacopa australis  flowers; photo: S.L. Winterton

Bacopa australis flowers; photo: S.L. Winterton

  Bacopa caroliniana , emersed; photo: S.L. Winterton

Bacopa caroliniana, emersed; photo: S.L. Winterton

  Bacopa caroliniana , submersed; photo: S.L. Winterton

Bacopa caroliniana, submersed; photo: S.L. Winterton

  Bacopa caroliniana  flower; photo: S.L. Winterton

Bacopa caroliniana flower; photo: S.L. Winterton

  Bacopa lanigera , emersed; photo: S.L. Winterton

Bacopa lanigera, emersed; photo: S.L. Winterton

  Bacopa lanigera  flower; photo: S.L. Winterton

Bacopa lanigera flower; photo: S.L. Winterton

  Bacopa monnieri , emersed; photo: S.L. Winterton

Bacopa monnieri, emersed; photo: S.L. Winterton

  Bacopa monnieri  flower; photo: S.L. Winterton

Bacopa monnieri flower; photo: S.L. Winterton

  Bacopa monnieri  flower; photo: S.L. Winterton

Bacopa monnieri flower; photo: S.L. Winterton

  Bacopa myriophylloides , emersed; photo: S.L. Winterton

Bacopa myriophylloides, emersed; photo: S.L. Winterton

  Bacopa myriophylloides , emersed; photo: S.L. Winterton

Bacopa myriophylloides, emersed; photo: S.L. Winterton

  Bacopa myriophylloides , submersed; photo: S.L. Winterton

Bacopa myriophylloides, submersed; photo: S.L. Winterton

  Bacopa myriophylloides  flower; photo: S.L. Winterton

Bacopa myriophylloides flower; photo: S.L. Winterton

  Bacopa salzmannii ; photo: S.L. Winterton

Bacopa salzmannii; photo: S.L. Winterton

  Bacopa salzmannii  flower; photo: S.L. Winterton

Bacopa salzmannii flower; photo: S.L. Winterton

  Bacopa serpyllifolia , emersed; photo: S.L. Winterton

Bacopa serpyllifolia, emersed; photo: S.L. Winterton

  Bacopa serpyllifolia  flower; photo: S.L. Winterton

Bacopa serpyllifolia flower; photo: S.L. Winterton